Fungalpedia – Note 1788, Pseudombrophilaceae
Pseudombrophilaceae Ekanayaka, K.D. Hyde, Q. Zhao and E.B.G. Jones
Index Fungorum, Facesoffungi, MycoBank, GenBank.
Classification: Incertae sedis, Incertae sedis, Incertae sedis, Pezizomycetes, Pezizomycotina, Ascomycota, Fungi
Ascomata apothecial or cleistothecial. Ectal excipulum omposed of textura angularis to globulosa or irregularly lobed cells, excipular hair present. Medullary excipulum composed of loosely arranged hyphae, sometimes unseparable from ectal excipulum. Asci operculate or inoperculate and non-amyloid. Ascospores globose to ellipsoid, guttulate. Paraphyses present or absent, filiform when present.
Notes: Heydenia, Lasiobolidium and Orbicula produce cleistothecial ascomata (Hansen et al. 2005; Doveri et al. 2013) while Pseudombrophila has apothecia. Lasiobolidium and Orbicula were initially placed within Eoterfeziaceae (Malloch and Cain 1971; Malloch and Benny 1973; Benny and Kimbrough 1980) and later transferred to Pyronemataceae (Hansen et al. 2005; Hansen and Pfister 2006; Perry et al. 2007). Hansen et al. (2005) showed that the genera Lasiobolidium and Orbicula are phylogenetically close to Pseudombrophila while Leuchtmann and Cle´menc¸on (2012) showed that Heydenia, Orbicula, Pseudombrophila and Lasiobolidium are monophyletic. Jaklitsch et al. (2016) suggested these four genera formed an independent lineage.
Type genus: Pseudombrophila deerrata (P. Karst.) Seaver, North American Cup-fungi, (Operculates) (New York): 141 (1928).