Fungalpedia – Note 3068, Porotheleaceae

 

Porotheleaceae Murrill.

Citation when using this data: Senanayake IC et al. 2023 – Fungalpedia.

Index Fungorum, Faceoffungi, MycoBankGenBank.

Classification: AgaricalesAgaricomycetidaeAgaricomycetesAgaricomycotinaBasidiomycota, Fungi.

The family Porotheleaceae was phylogenetically revised and 14 genera were accepted (Antonín et al. 2019He et al. 2019Vizzini et al. 2019; Kalichman et al. 2020; Matheny et al. 2020). These studies indicated that taxonomic placements of some genera are still doubtful. Consiglio et al. (2022) mentioned that there are no obvious morphological or ecological synapomorphies shared by all members of the pale-spored Porotheleaceae s. stricto, with the exception of their lignicolous habit and the presence of clamp connections. Unfortunately, the same situation within the genus Porotheleum is amended by Consiglio et al. (2022). Moreover, their phylogenetic trees (Consiglio et al. 2022, p. 104, 106 and 107), as well as our tree (Fig. 152; Senanayake IC et al. 2023) support that Porotheleum sensu Consiglio et al. contains more genera. All specimens classified by Consiglio et al. (2022) under the genus Porotheleum formed a clade with ML/BI = 72%/0.95 statistical support (Fig. 154; Senanayake IC et al. 2023). We identified several well-separated lineages as, 1) Clitocybula albida (P. albidum), 2) P. dominingense and P. nigripes 3) P. fimbriatum and Porotheleum sp. G0006 4) P. parvulum, 5) Porotheleum sp. PBM4282, and a lineage of 6) Porotheleum omphaliiforme and our new collections (BRNM 552658, BRNM 552721, BRNM 552654). 

Porotheleum fimbriatum (Pers.) Fr. is the type species of Porotheleum Fr. Thus, lineage 3) P. fimbriatum and Porotheleum sp. G0006 represents Porotheleum sensu stricto. Clitocybula albida A.K. Dutta, K. Acharya & Antonín clustered with Porotheleum in the phylogenetic analysis of Consiglio et al. (2022). Phylogenetic analysis in this study (Fig.152; Senanayake IC et al. 2023) also showed that, Clitocybula albida and Porotheleum albidum form a separate clade and however, further studies are required to confirm its generic position using additional materials and more genes, since only ITS data for C. albida are available (Dutta et al. 2018). Porotheleum dominingense and P. nigripes form well-separated lineage and clearly represent a new genus, described here as Vizzinia.

Type species: Porotheleum Fr.

Other accepted species: Species Fungorum – search Porotheleaceae.

 

References

Antonín V, Borovička J, Holec J, Piltaver A, Kolařík M. 2019 – Taxonomic update of Clitocybula sensu lato with a new generic classification. Fungal Biol 123: 431–447.

Consiglio G, Vizzini A, Cooper J, Marchetti M et al. 2022 – The agaricoid members of the genus Porotheleum (Porotheleaceae, Agaricales), Porotheleum emend., Porotheleaceae s. stricto, and new genera for Agaricus floccipes and Mycena subalpine. Riv Micol 64(2): 152–179.

Dutta AK, AntonínV, Barui R, Acharya K. 2018 – A new species of Clitocybula (Marasmiaceae) from West Bengal, India. Nova Hedwigia 107(1): 195–203.

He MQ, Zhao RL, Hyde KD, Begerow D et al. 2019 – Notes, outline, and divergence times of Basidiomycota. Fungal Divers 99: 105–367.

Matheny PB, Hobbs AM, Esteve-Raventós F. 2020 – Genera of Inocybaceae: New skin for the old ceremony. Mycologia 112: 83–120.

Senanayake IC, Rossi W, Leonardi M. et al. 2023 – Fungal diversity notes 1611–1716: taxonomic and phylogenetic contributions on fungal genera and species emphasis in south China. Fungal Diversity 122, 161–403.

Vizzini A, Picillo B, Perrone L, Dovana F. 2019 – Chrysomycena perplexa gen. et sp. nov. (Agaricales, Porotheleaceae), a new entity from the Lazio region. Riv Micol Romana 107: 96–107. 

 

Entry by

Indunil C. Senanayake, Innovative Institute for Plant Health/ Key Laboratory of Green Prevention and Control on Fruits and Vegetables in South China, Ministry of Agriculture and Rural Affairs, Zhongkai University of Agriculture and Engineering, Haizhu District, Guangzhou 510225, Guangdong, China., Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand., Guangdong Provincial Key Laboratory for Plant Epigenetics, College of Life Science and Oceanography, Shenzhen University, 3688, Nanhai Avenue, Nanshan, Shenzhen, Guangdong 518055, China., University of L’Aquila, Department MeSVA, sect. Environmental Sciences, Via Vetoio, 67100 Coppito (AQ), Italy.

 

Published online 31 August 2026