Fungalpedia – Note 738, Astrosphaeriellaceae

 

Astrosphaeriellaceae Phookamsak & K.D. Hyde

Citation when using this data: Karimi O et al. 2025 (in prep.) – Fungalpedia, Ascomycota.

Index FungorumFacesoffungiMycoBankGenBank

Classification: PleosporalesPleosporomycetidaeDothideomycetesPezizomycotinaAscomycota, Fungi

Astrosphaeriellaceae was introduced by Phookamsak et al. (2015) within Pleosporales to include Astrosphaeriella and Pteridiospora, based on both morphological features and combined phylogenetic analyses of LSU, SSU, and tef-1α. Since then, several additional genera have been incorporated into the family (Liu et al. 2018Wanasinghe et al. 2018Wijayawardene et al. 2018Jayasiri et al. 2019Dong et al. 2020Konta et al. 2023). Currently, 14 genera are recognized within Astrosphaeriellaceae (Hyde et al. 2024Zhang et al. 2024a). Members of Astrosphaeriellaceae have been reported as saprobic or parasitic on palms, bamboo, Quercus species, or robust grasses (Zhang et al. 2024a). The sexual morph is characterized by uni-loculate, solitary to gregarious, erumpent to superficial, glabrous, brittle, and carbonaceous ascomata. These structures are dark opaque, conical or mammiform, thick-walled with uneven thickness, and poorly developed at the base. They may be surrounded by ruptured, reflexed, stellate host tissue remnants at the base. The ascomata are composed of thick, opaque, and melanized cells, with palisade-like cells present at the rim of the peridium. In the hamathecium, dense, anastomosing, trabeculate pseudoparaphyses are observed. The asci are 8-spored, bitunicate, fissitunicate, cylindrical to cylindric-clavate, pedicellate, and rounded apically with an ocular chamber or a J-, subapical ring. The ascospores are subfusoid to fusiform, obclavate to ellipsoidal, or limoniform, hyaline or pale brown to reddish brown, septate, constricted at the septum, and smooth-walled. Appendages and a mucilaginous sheath may be present on the ascospores. The asexual morph is reported as coelomycetous or hyphomycetous (Hongsanan et al. 2020). 

Type genus: Astrosphaeriella Syd. & P. Syd., Annls mycol. 11(3): 260 (1913).

 

References

Dong W, Wang B, Hyde KD, McKenzie EHC, et al. 2020 – Freshwater Dothideomycetes. Fungal Diversity 105, 319–575. 

Hongsanan S, Hyde KD, Phookamsak R, Wanasinghe DN, et al. 2020 – Refined families of Dothideomycetes: Dothideomycetidae and Pleosporomycetidae. Mycosphere 11, 1553–2107.

Hyde KD, Noorabadi MT, Thiyagaraja V, He MQ, et al. 2024 – The 2024 Outline of Fungi and fungus-like taxa. Mycosphere 15, 5146–6239. 

Jayasiri SC, Hyde KD, Jones EBG, McKenzie EHC et al. 2019 – Diversity, morphology and molecular phylogeny of Dothideomycetes on decaying wild seed pods and fruits. Mycosphere 10, 1–186. 

Konta S, Tibpromma S, Karunarathna SC, Samarakoon MC et al. 2023 – Morphology and multigene phylogeny reveal ten novel taxa in Ascomycota from terrestrial palm substrates (Arecaceae) in Thailand. Mycosphere 14, 107–152.

Liu LL, Long QD, Kang JC, Zhang X et al. 2018 – Morphology and phylogeny of Mycopepon. Mycosphere 9, 779–789.

Phookamsak R, Norphanphoun C, Tanaka K, Dai DQ et al. 2015 – Towards a natural classification of Astrosphaeriella-like species; introducing Astrosphaeriellaceae and Pseudoastrosphaeriellaceae fam. nov. and Astrosphaeriellopsis, gen. nov. Fungal Diversity 74, 143–197.

Wanasinghe DN, Jeewon R, Jones EBG, Boonmee S, et al. 2018 – Novel palmicolous taxa within Pleosporales: multigene phylogeny and taxonomic circumscription. Mycological Progress 17, 571–590. 

Wijayawardene NN, Hyde KD, Lumbsch HT, Liu JK, et al. 2018 – Outline of Ascomycota: 2017. Fungal Diversity 88, 167–263. 

Zhang SN, Hyde KD, Jones EG, Yu XD et al. 2024a – Current insights into palm fungi with emphasis on taxonomy and phylogeny. Fungal Diversity 127, 55–301.

 

Entry by

Omid Karimi, State Key Laboratory of Discovery and Utilization of Functional Components in Traditional Chinese Medicine & School of Pharmaceutical Sciences, Guizhou Medical University, Guian New District, Guizhou 550004, China, Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand, School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand

 

Published online 28 July 2025